1. Key Laboratory of Wetland Ecology and Environment, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China
2. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China
3. USDA, Agricultural Research Service, San Joaquin Valley Agricultural Science Center, 9611 South Riverbend Avenue, Parlier, CA 93648-9757, USA
4. Department of Natural Sciences, Middlesex University, Hendon, London NW4 4BT, UK