1. School of Marine Science and Engineering, Qingdao Agricultural University, Qingdao 266000, China
2. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266000, China
3. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266000, China
4. Rizhao Duobao Aquaculture Co., Ltd., Rizhao 276800, China