1. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, China
2. Shandong Provincial Center of Forest and Grass Germplasm Resources, Institute of Resource Cultivation, Jinan 250102, China
3. Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, China