1. Key Laboratory of Ecological Safety and Sustainable Development in Arid Lands, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
2. Shapotou Desert Research and Experiment Station, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
3. University of Chinese Academy of Sciences, Beijing 101408, China
4. Institute of Desertification Control, Ningxia Academy of Agricultural and Forestry Sciences, Yinchuan 750002, China
5. College of Forestry, Gansu Agricultural University, Lanzhou 730070, China