1. Engineering Research Center of Agricultural Microbiology Technology, Ministry of Education, Heilongjiang University, Harbin 150500, China
2. Heilongjiang Province Key Laboratory of Ecological Restoration and Resource Utilization for Cold Region, School of Life Sciences, Heilongjiang University, Harbin 150080, China
3. Institute of Nature and Ecology, Heilongjiang Academy of Sciences, Harbin 150040, China