1. Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
2. Department of Microbiology and Immunology, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
3. Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
4. Institute for the Oceans and Fisheries, University of British Columbia, Vancouver, BC V6T 1Z4, Canada