1. Department of Architecture, University of Pretoria, Pretoria 0002, South Africa
2. Department of Biology, Aarhus University, DK-8000 Aarhus C, Denmark
3. Center for Ecological Dynamics in a Novel Biosphere (ECONOVO) & Center for Biodiversity Dynamics in a Changing World (BIOCHANGE), Department of Biology, Aarhus University, DK-8000 Aarhus C, Denmark