1. Department of Infectious Diseases, Melbourne Medical School, The University of Melbourne, The Peter Doherty Institute for Infection and Immunity, Melbourne, VIC 3000, Australia
2. Victorian Infectious Diseases Reference Laboratory, Royal Melbourne Hospital, The Peter Doherty Institute for Infection and Immunity, Melbourne, VIC 3000, Australia
3. Curtin Medical School, Curtin University, Perth, WA 6102, Australia