1. Wildlife Conservation Research Unit, Department of Biology, University of Oxford, The Recanati-Kaplan Centre, Tubney House, Abingdon Road, Tubney, Abingdon OX13 5QL, UK
2. Department of Chemistry and Bioscience, Aalborg University, Fredrik Bajers Vej 7H, DK-9220 Aalborg, Denmark
3. Naturama, 30 Dronningemaen, DK-5700 Svendborg, Denmark
4. Department of Biology, University of Southern Denmark, 55 Campusvej, DK-5230 Odense M, Denmark
5. Department of Geosciences and Natural Resource Management, Section for Forest, Nature and Biomass, Copenhagen University, 23 Rolighedsvej, DK-1958 Frederiksberg C, Denmark
6. Interdisciplinary Centre on Population Dynamics (CPop), University of Southern Denmark, 55 Campusvej, DK-5230 Odense M, Denmark