1. Colin Ratledge Center for Microbial Lipids, School of Agricultural Engineering and Food Science, Shandong University of Technology, Zibo 255049, China
2. School of Biology and Brewing Engineering, Taishan University, Taian 271000, China
3. Department of Botany and Microbiology, Faculty of Science, Al-Azhar University, Assiut 71524, Egypt