1. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. College of Life and Environmental Sciences, Minzu University of China, Beijing 100081, China
4. Herbarium Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Cibinong, Bogor 16911, Indonesia