1. Key Laboratory of Forest Ecology in Tibet Plateau, Ministry of Education, Institute of Tibet Plateau Ecology, Tibet Agricultual & Animal Husbandry University, Nyingchi 860000, China
2. National Key Station of Field Scientific Observation & Experiment, Nyingchi 860000, China
3. Key Laboratory of Biodiversity Conservation of National Forestry and Grassland Administration, Ecology and Nature Conservation Institute, Chinese Academy of Forestry, Beijing 100091, China