1. Department of Medicine, University of Cambridge, Cambridge CB2 0QQ, UK
2. Department of Pathology, University of Cambridge, Cambridge CB2 0QQ, UK
3. Infection Biology, Global Center for Pathogen and Human Health Research, Lerner Research Institute, Cleveland Clinic, Cleveland, OH 44196, USA
4. University of Sao Paulo, Sao Paulo 05508-060, Brazil
5. Infection and Immunity, University College London, London WC1E 6BT, UK