1. Institute of Eastern-Himalaya Biodiversity Research, Dali University, Dali 671003, China
2. Collaborative Innovation Center for Biodiversity and Conservation in the Three Parallel Rivers Region of China, Dali 671003, China
3. The Provincial Innovation Team of Biodiversity Conservation and Utility of the Three Parallel Rivers Region, Dali University, Dali 671003, China
4. State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China
5. Yunling Black-and-White Snub-Nosed Monkey Observation and Research Station of Yunnan Province, Dali 671003, China