1. CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China
2. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China
3. E’erguna Wetland Ecosystem National Research Station, Hulunbuir 022250, China
4. School of Environment, Liaoning University, Shenyang 110036, China
5. Erguna Forest-Steppe Ecotone Research Station, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China