1. Key Laboratory of National Forestry and Grassland Administration of Karst Mountain Biodiversity Conservation in Southwest China of Guizhou Province, Guizhou Academy of Forestry, Guiyang 550005, China
2. The National Positioning Observation and Research Station of Leigong Mountain Forest Ecosystem of Guizhou Province, Kaili 557100, China
3. Guizhou Forest Investigation and Planning Institute, Guiyang 550003, China