Effects of Neighborhood Competition and Stand Structure on the Productivity of Pure and Mixed Larix principis-rupprechtii Forests

Abstract

Understanding the factors influencing tree productivity is central to forest ecology. However, the relative contributions of neighborhood interactions, tree species diversity, and tree size to larch (Larix principis-rupprechtii) productivity require further study. Three plots in the Guandi Mountains, Shanxi Province, were set up for each of the following forest types: natural pure larch forest (PL), mixed larch and birch (Betula platyphylla) forest (LB), and mixed larch and spruce (Picea asperata) forest (LS). Based on the tree size-stratified sampling method, a total of 318 tree core samples were collected. A linear mixed model was used to analyze the effects of tree size, dominance, mixing, and neighborhood competition on larch productivity. Birch and spruce promoted larch growth at the stand and individual tree levels, and birch exhibited a more significant facilitating effect. Intraspecific competition was the main factor affecting larch growth. When the intensity of competition among trees was low, the basal area increment (BAI) of larch in the mixed forests was higher than that in the pure forest. However, with increasing competition, the BAI of larch was lower in the mixed forests than in the pure forest. Factors including tree size, dominance, and mingling were positively correlated with the BAI of larch. With increasing tree size, the BAI of larch was higher in the mixed forests than in the pure forest and higher in LB than in LS. When the dominance was less than 0.5, the BAI of larch was higher in the pure forest than in the mixed forests and higher in LS than in LB. With increasing dominance, the BAI of larch was higher in the mixed forests than in the pure forest. The BAI of larch increased with an increasing mixing degree in the mixed forests, and the increasing trend of BAI was larger in LB than in LS. Larch productivity was influenced mainly by neighborhood interactions and stand structure. Improving neighborhood tree diversity and increasing the large tree proportion and dominance of larch will be helpful for improving larch productivity in mixed forests.