1. State Key Laboratory of Herbage Improvement and Grassland Agro-Ecosystems, Lanzhou University, Lanzhou 730000, China
2. Heshui Branch of Ziwuling Forestry Administration of Gansu Province, Qingyang 745400, China
3. Key Laboratory for Earth Surface Processes of the Ministry of Education, Institute of Ecology, College of Urban and Environmental Sciences, Peking University, Beijing 100871, China