1. College of Life Sciences, Southwest Forestry University, Kunming 650224, China
2. Key Laboratory for Forest Genetic and Tree Improvement and Propagation in University of YunnanProvince, Southwest Forestry University, Kunming 650224, China
3. College of Forestry, Southwest Forestry University, Kunming 650224, China
4. Key Laboratory of Biodiversity Conservation in Southwest China, State Forestry Administration, Southwest Forestry University, Kunming 650224, China