1. Hainan Yazhou Bay Seed Laboratory, Sanya Nanfan Research Institute of Hainan University, Hainan University, Sanya 572025, China
2. College of Tropical Agricultural and Forestry, Hainan University, Haikou 570228, China
3. Queensland Alliance for Agriculture and Food Innovation, The University of Queensland, Health and Food Sciences Precinct, Coopers Plains, QLD 4108, Australia