1. Watershed Hydrology and Ecology Research Division, Environment and Climate Change Canada, University of Victoria Queenswood Campus, 2474 Arbutus Road, Victoria, BC V8N 1V8, Canada
2. Department of Geography, University of Victoria, Victoria, BC V8W 2Y2, Canada
3. Department of Civil Engineering, University of Victoria, Victoria, BC V8W 2Y2, Canada
4. Environment and Climate Change Canada, Canadian Rivers Institute, Faculty of Forestry and Environmental Management, University of New Brunswick, Fredericton, NB E3B 5A3, Canada
5. Environment and Climate Change Canada, Canadian Rivers Institute, Department of Biology, University of New Brunswick, Fredericton, NB E3B 5A3, Canada