1. Department of Microbiology, Graduate School, Daejeon University, Daejeon 34520, Republic of Korea
2. Department of Biomedical Laboratory Science, Daejeon University, Daejeon 34520, Republic of Korea
3. Soonchunhyang Institute of Medi-Bio Science, Soonchunhyang University, Cheonan 31151, Republic of Korea
4. Department of Bacteriology, University of Wisconsin-Madison, Madison, WI 53706, USA