1. College of Life Sciences and Oceanography, Shenzhen University, Shenzhen 518060, China
2. South China Sea Sea Area and Island Center, Ministry of Natural Resources, Guangzhou 510301, China
3. Key Laboratory of Marine Environmental Survey Technology and Application, Ministry of Natural Resources, Guangzhou 510301, China
4. College of Marine Science and Technology, China University of Geosciences, Wuhan 430079, China