1. Institute of Marine Biological Resources and Inland Waters, Hellenic Centre for Marine Research, 19013 Anavissos, Greece
2. Bristol Zoological Society, Bristol Zoo Gardens, Bristol BS8 3HA, UK
3. Centre for Research in Biosciences, University of the West of England, Coldharbour Lane, Bristol BS16 9QY, UK
4. Wildfowl & Wetlands Trust (WWT), Slimbridge, Gloucester GL2 7BT, UK