1. Institute of Horticulture, Guizhou Academy of Agricultural Sciences/Guizhou Horticultural Engineering Technology Research Center, Guiyang 550006, China
2. Key Laboratory of Crop Gene Resources and Germplasm Innovation in Karst Mountain Area of Agriculture and Rural Ministry, Guiyang 550006, China
3. State Key Laboratory of Crop Genetics & Germplasm Enhancement and Utilization, Ministry of Agriculture and Rural Affairs Key Laboratory of Biology and Germplasm Enhancement of Horticultural Crops in East China, College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China
4. College of Horticulture, Sichuan Agricultural University, Chengdu 611130, China
5. College of Biology and Agricultural Technology, Zunyi Normal College, Zunyi 563006, China