1. Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin 541006, China
2. College of Life Sciences, Guangxi Normal University, Guilin 541006, China
3. School of Pharmacy, Guilin Medical University, Guilin 541006, China