1. Doctoral School of Natural Ecosystems, University of Mahajanga, Mahajanga 401, Madagascar
2. Sciences de la Vie et de l’Environnement, Faculté des Sciences, de Technologies et de l’Environnement, University of Mahajanga, BP 652, Mahajanga 401, Madagascar
3. American Museum of Natural History, Department of Herpetology, New York, NY 10024, USA