1. School of Agriculture, Yunnan University, Kunming, Yunnan 650504, China,
2. Department of Ecology and Evolutionary Biology, Yale University, P.O. Box 208106, New Haven, Connecticut 06520, USA,
3. Institute of Ecology and Geobotany, School of Ecology and Environmental Sciences, Yunnan University, Kunming, Yunnan 650091, China,
4. Institute of Ecology and Geobotany, School of Ecology and Environmental Sciences & Yunnan Key Laboratory for Plateau Mountain Ecology and Restoration of Degraded Environments, Yunnan University, Kunming, Yunnan 650091, China,