1. Professor, Graduate School of Environmental and Life Science, Okayama Univ., 3-1-1 Tsushima-naka, Kita-ku, Okayama 700-8530, Japan (corresponding author).
2. Assistant Professor, Graduate School of Environmental and Life Science, Okayama Univ., 3-1-1 Tsushima-naka, Kita-ku, Okayama 700-8530, Japan.
3. Lecturer, Graduate School of Agriculture, Kyoto Univ., Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto 606-8502, Japan.