1. Medical Research Council Centre for Reproductive Health (D.R., J.W., L.C., S.E.S., L.M., A.F., R.T.M., L.B.S.), University of Edinburgh, The Queen's Medical Research Institute, Edinburgh EH16 4TJ, United Kingdom
2. University/BHF Centre for Cardiovascular Science (J.W., P.W.F.H.), University of Edinburgh, The Queen's Medical Research Institute, Edinburgh EH16 4TJ, United Kingdom
3. Department of Orthopaedics (R.J.W.), University of Edinburgh, Edinburgh Eh16 4SB, United Kingdom;
4. Clinical Research Imaging Centre (C.D.G.), University of Edinburgh, The Queen's Medical Research Institute, Edinburgh EH16 4TJ, United Kingdom;
5. Institute of Biodiversity, Animal Health, and Comparative Medicine (P.J.O.), University of Glasgow, Garscube Campus, Glasgow G61 1QH, United Kingdom