1. Central Washington University, Biological Sciences , Ellensburg, WA 98926 , USA
2. Center for Reproductive Biology, Washington State University , Pullman, WA 99164 , USA
3. Reproductive & Developmental Biology Laboratory, National Institute of Environmental Health Sciences, National Institutes of Health, Research Triangle Park , NC 27709 , USA
4. Biological Sciences Division, University of Chicago Pritzker School of Medicine , Chicago, IL 60637 , USA