1. Makino Herbarium, Tokyo Metropolitan University, 1-1 Minami-Osawa, Hachioji, Tokyo 192-0397, Japan
2. Present address; The Kochi Prefectural Makino Botanical Garden 4200-6 Godaisan, Kochi Pref. 781-8125, Japan
3. Department of Botany, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan.
4. Department of Biology, Graduate School of Science, Chiba University, 1-33 Yayoi-cho, Inage, Chiba 263-8522, Japan