1. Institute of Biology; Freie Universität Berlin; Altensteinstr. 6 14195 Berlin Germany
2. Berlin-Brandenburg Institute of Advanced Biodiversity Research (BBIB); 14195 Berlin Germany
3. Centre d'Ecologie Fonctionnelle et Evolutive (CEFE); UMR 5175; CNRS, Université de Montpellier, Université Paul-Valéry Montpellier, EPHE; 1919 route de Mende 34293 Montpellier Cedex 5 France
4. School of Biological Sciences; University of California; Irvine California 92697 USA