1. Laboratoire de Biométrie et Biologie Evolutive, UMR 5558, CNRS Université Claude Bernard Lyon 1 Villeurbanne Cedex France
2. Department of Aquaculture and Fish Biology Hólar University Háeyri 1 Sauðárkrókur 550 Iceland
3. Department of Biology University of Southern Denmark Campusvej 55 Odense M 5230 Denmark
4. Department of Zoology University of Oxford Oxford OX1 3PS UK
5. Department of Biology Stanford University Stanford California 94305 USA