1. USGS Eastern Ecological Science Center SO Conte Anadromous Fish Laboratory 1 Migratory Way Turners Falls Massachusetts 01376 USA
2. Department of Environmental Conservation University of Massachusetts 160 Holdsworth Way Amherst Massachusetts 01003 USA
3. Department of Biology Monmouth University 400 Cedar Avenue West Long Branch New Jersey 07764 USA
4. Centre for Research into Ecological & Environmental Modelling, The Observatory, Buchanan Gardens University of St Andrews St Andrews Fife KY16 9LZ United Kingdom