1. Department of Environmental Conservation; University of Massachusetts; Amherst MA 01003 USA
2. Smithsonian Conservation Biology Institute; National Zoological Park; 1500 Remount Road Front Royal VA 22630 USA
3. Department of Biology; University of Maryland; College Park MD 20742 USA
4. Biodiversity and Climate Research Centre (BiK-F); Senckenberg Gesellschaft für Naturforschung, and Goethe Universität Frankfurt; Senckenberganlage, 25 60325 Frankfurt (Main) Germany
5. Wildlife Conservation Society; 2300 S Boulevard Bronx NY 10460 USA