1. The Hakubi Center for Advanced Research Kyoto University Kyoto Japan
2. Laboratory of Animal Ecology, Department of Zoology, Graduate School of Science Kyoto University Kyoto Japan
3. Center for Data Assimilation Research and Applications, Joint Support Center for Data Science Research Research Organization of Information and Systems Tokyo Japan
4. Regional Environment Conservation Division National Institute for Environmental Studies Ibaraki Japan
5. Federal Scientific Center of the East Asia Terrestrial Biodiversity Far Eastern Branch of the Russian Academy of Sciences Vladivostok Russia