1. Key Laboratory of Biodiversity and Biogeography; Kunming Institute of Botany; Chinese Academy of Sciences; Heilongtan, 132 Lanhei Road, Kunming Yunnan Province 650204 China
2. United States National Herbarium; Department of Botany; Smithsonian Institution; National Museum of Natural History; MRC-166, P.O. Box 37012 Washington, D.C. 20013-7012 U.S.A.