1. School of Anatomical Sciences, Faculty of Health Sciences University of the Witwatersrand Johannesburg Republic of South Africa
2. Department of Anthropology and Center for the Advanced Study of Human Paleobiology The George Washington University Washington District of Columbia USA
3. Borås Zoo Borås Sweden
4. Centre for Zoo and Wild Animal Health, Copenhagen Zoo Frederiksberg Denmark
5. Department of Anatomy and Neurobiology Boston University, School of Medicine Boston Massachusetts USA
6. Department of Physiology, Anatomy and Genetics University of Oxford Oxford UK