1. Institute of Ecology and Environmental Sciences of Paris; UMR 7618; UPMC-CNRS; Paris 75005 France
2. Department of Infectious Diseases Epidemiology; University of London Imperial College of Science, Technology & Medicine; Norfolk Place, W21PG London UK
3. UMR 5558 Laboratoire Biométrie et Biologie Evolutive; Université Lyon 1 - CNRS; Villeurbanne F-69622 France
4. INRA - CNRS - Université Nice Sophia Antipolis; UMR 1355 - 7254 Institut Sophia Agrobiotech; Sophia Antipolis 06903 France
5. AgroParisTech; Paris F-75005 France