1. Department of Biological SciencesUniversity of Cape Town Cape Town South Africa
2. School of Earth SciencesUniversity of Bristol Bristol UK
3. Groningen Biomolecular Sciences and Biotechnology Institute and Zernike Institute for Advanced MaterialsUniversity of Groningen Groningen The Netherlands
4. UMR 7205 Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum National d'Histoire Naturelle, CNRS, Sorbonne UniversitéEPHE Paris France