1. U.S. Geological Survey Pacific Island Ecosystems Research Center Box 44, Hawaiʻi National Park HI 96718 USA
2. Hawaiʻi Cooperative Studies Unit University of Hawaiʻi Hilo PO Box 44, Hawaiʻi National Park HI 96718 USA
3. Department of Biology Tufts University 200 College Avenue Medford MA 02155 USA
4. Pacific Islands National Wildlife Refuge Complex U.S. Fish and Wildlife Service Honolulu HI 96850 USA