1. Department of Neurobiology; David Geffen School of Medicine at UCLA, University of California; Los Angeles California 90095
2. Department of Medicine; David Geffen School of Medicine at UCLA, University of California; Los Angeles California 90095
3. Jules Stein Eye Institute, David Geffen School of Medicine at UCLA, University of California; Los Angeles California 90095
4. Physiology; Genetics and Microbiology; University of Alicante; 03690 Alicante Spain
5. CURE-Digestive Diseases Research Center, David Geffen School of Medicine at UCLA, University of California; Los Angeles California 90095
6. Veterans Administration Greater Los Angeles Healthcare System; Los Angeles California 90073