1. Arthritis Research UK Centre for Osteoarthritis Pathogenesis, Kennedy Institute of Rheumatology, University of Oxford; Oxford UK
2. Fortius Clinic, Imperial College Healthcare NHS Trust, St. Mary's Hospital; London UK
3. NIHR Musculoskeletal Biomedical Research Unit, University of Oxford, Oxford, UK, and University of Southampton; Southampton UK
4. Arthritis Research UK Centre for Osteoarthritis Pathogenesis, Kennedy Institute of Rheumatology, University of Oxford, Oxford, UK, and Fortius Clinic; London UK