1. Department of Agricultural, Food and Environmental Science; University of Perugia; Via San Costanzo 06126 Perugia Italy
2. Department of Agriculture, Environmental and Food Science, and Industrial Yeasts Collection DBVPG; University of Perugia; Borgo XX Giugno, 74 06121 Perugia Italy
3. Italian Brewing Research Centre; University of Perugia; Via San Costanzo 06126 Perugia Italy