1. Department of Geography National Taiwan University Taipei 10617 Taiwan
2. Research Center for Future Earth National Taiwan University Taipei 10617 Taiwan
3. Department of Ecology and Evolutionary Biology University of Arizona Tucson Arizona 85721 USA
4. Earth & Environment Boston University Boston Massachusetts 02215 USA
5. Department of Biology University of Maryland College Park Maryland 20742 USA
6. The Santa Fe Institute Santa Fe New Mexico 87501 USA