1. Centre for Sport, Exercise and Osteoarthritis Research Versus Arthritis, School of Life Sciences The University of Nottingham Nottingham UK
2. National Institute for Health and Care Research (NIHR) Nottingham Biomedical Research Centre Nottingham University Hospitals NHS Trust and University of Nottingham Nottingham UK
3. MRC/Versus Arthritis Centre for Musculoskeletal Ageing Research, Schools of Life Sciences and Medicine University of Nottingham Nottingham UK
4. Sport and Health Sciences The University of Exeter Exeter UK
5. Ohio Musculoskeletal and Neurological Institute, Heritage College of Osteopathic Medicine Ohio University Athens OH USA
6. Department of Biomedical Sciences University of Padua Padua Italy
7. Core Facility for Integrated Microscopy The University of Copenhagen Copenhagen Denmark