1. School of Aquatic and Fishery Sciences University of Washington Box 355020 Seattle Washington 98195 USA
2. National Center for Ecological Analysis and Synthesis University of California, Santa Barbara 735 State St. #300 Santa Barbara California 93101 USA
3. Bren School of Environmental Science and Management University of California, Santa Barbara Santa Barbara California 93106 USA
4. Imperial College London Silwood Park Campus, Buckhurst Rd. Ascot SL57PY UK