1. Department of Biotechnology Graduate School of Agriculture and Life Sciences The University of Tokyo 1-1-1 Yayoi Bunkyo-ku Tokyo 113-8657 Japan
2. Noda Institute for Scientific Research 338 Noda, Noda City Chiba 278-0037 Japan
3. Collaborative Research Institute for Innovative Microbiology Graduate School of Agriculture and Life Sciences The University of Tokyo 1-1-1 Yayoi Bunkyo-ku Tokyo 113-8657 Japan