1. Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research; Chinese Academy of Sciences; Beijing China
2. Anhui Agricultural University; Hefei China
3. Nicholas School of the Environment; Duke University; Durham North Carolina USA
4. State Key Laboratory of Forest and Soil Ecology, Institute of Applied Ecology; Chinese Academy of Sciences; Shenyang China
5. Department of Biology and Graduate Degree Program in Ecology; Colorado State University; Fort Collins Colorado USA